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acts of urination adult male dogs alternate days amount of testosterone androgen Animal 3b Animal 5a animal elevated animal urinated anosmic assume the puppy castrated animal codliver oil completely individuated control littermates copulatory activity copulatory behavior days old delay animal delay cage digitograde elevate the leg elevation behavior estrogens exercise yard female puppies female rats female sex hormones frequently gross torsion movement hindquarters hormone activity hormone injections injection of female injection of testosterone last injection leg during urination leg when urinating leg-elevation articulated leg-elevation behavior leg-elevation pattern leg-raising male copulatory pattern male leg-elevation male puppies male sex hormone maternal behavior maturation micturition neural organization normal adult male normal female olfaction olfactory bulbs ovarectomized Psychol puppy posture puppy stance puppy urination posture puppy urination stance reversion sensori-motor mechanism sling testicular testosterone propionate theelin trial and error urine was expelled vitamin B1 weeks old young puppy