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abundance Agua Hedionda lagoon Allometric Anacapa Island analysis April Aristotle's lantern benthic Biol body wall calcein City of San CN CN collected from Agua components covariance demipyramids differ disease distribution Dry Weight Ebert Echinoderm echinoid Ecol ecotypic environmental equation estimates Figure Four Locations Frenchy's Cove genital plates gonad gonad weight Gonad Wet Weight grazing growth and mortality growth study growth zones habitats Hope Rock individuals inshore and offshore invertebrate jaw lengths July July 27 kelp bed Long Beach Lytechinus from Four Lytechinus pictus Macrocystis Macrocystis pyrifera Mean Test Diameter mortality coefficient nonlinear regression nutrient offshore populations outer lagoon outfall parameters Pearse phenotypic pictus pictus collected pictus from Agua Point Loma PX PX recruitment Relationship Between Test reproductive resource allocation San Diego sea urchin Southern California Bight spawning species specimens spicules spine Strongylocentrotus purpuratus tagged urchins Test Diameter test height Test Plate tetracycline transects transplanted trawling Walford plot