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abundance Albert Temple amount of testosterone annual cycle behavior bill color breeding season collection period correlated daylengths eggs environmental estradiol estrogen ethanol female Starlings female's Figure final maturation FSH and LH FSH release FSH secretion functional gonadal hypothalamus increase interstitial cells interstitial Leydig cells Kessel levels Leydig cells LH release LH secretion Lofts male Starlings midpoint ng/lOml ovarian follicles ovarian weight ovary oviduct ovulation peak photoperiods photorefractory photosensitive photosensitive period photostimulation pituitary gland pituitary gonadotropin content plasma plasma testosterone point is located points represent collections post-breeding prolactin psychic stimuli recovery refractory period represents the mean reproductive cycle reproductive organs reproductive system response result seasonal changes seasonal high values seasonal low Seasonal variation second broods seminal sacs seminiferous tubules single day spermatocytes spermatogonia spermatozoa Starling Sturnus vulgaris Starling's steroids testes testosterone production testosterone secretion testosterone titers thin layer plate Tienhoven Tixier-Vidal tube winter Witschi yearly cycle