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adjacent agricultural Amberley autumn burning Canterbury and Marlborough carried cattle cent chloropicrin Class I infestation cocksfoot control and eradication crop cultivation dense density deposited English grass pasture establishment of nassella examined experimental factor farmers favourable Fruit Content fruit dispersal fruit production fruits of nassella germinated fruits germination grassland grazing growth grubbing heavily infested heavy Hurunui River inches indicated indigenous tussocks infested area infested localities inflorescences instance investigation land layer Leithfield Leptospermum scoparium Lower Hutt lucerne mature tussocks method modified tussock-grassland moisture content mortality mother patch nassella tussock establishment nassella tussock fruits nassella tussock infestation nassella tussock seedlings non-infested North Canterbury north west slopes north-west number of fruits Omihi Valley ploughable ploughing present properties regeneration relatively river bed areas rolling country samples season seed sheep shown slopes soil moisture spread stage Stipa sward terraces unploughable vegetative cover Waikari Waipara River Wallaceville weed Weka wind-borne fruits