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abundance animals Ansell April areas ash-free attached autumn and winter Bayne Biol biomass bivalves collectors Conwy Dare Davies density deposit dry flesh weight early plantigrades Ecol Eider estimated Estuary Figure Fishery Invest flatfish Flesh ash-inclusive ground growth curve Heysham hydroids ICES CM intertidal intertidal mussels Irish Sea July June Knott End mussels Knott End Skear large numbers larvae late plantigrades live weight Lond low-water mark Main Skear maximum mean length Menai Straits Mercenaria mimeo mixed year-classes Morecambe Bay mortality rate mussel beds mussel populations mussel-mud mussels Mytilus edulis North Wales older mussels Overwintering plankton population density predation red algae Roosebeck salinity sampling Seasonal changes Seed September settlement occurred settlement of Mytilus shell growth shell weight Shellfish Comm similar Skear study plot South America spat spatfalls spring tides starfish storms sublittoral substrate summer survival temperatures Theisen tidal Walne winter plantigrades winter settlement year-class young mussels Ythan Estuary