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5-merous A. H. Moore Acad achenes Acmella acute to acuminate anthers apex acute apex rounded Argentina Arts 42 bellidioides biseriate brachyglossa broadly ovate Ceratocephalus Chromosome number ciliate with straight-tipped cm long corollas DeCandolle decumbens decumbent densely ciliate densely pilose dentate diameter diploid disc corollas Disc florets discoid filipes glabrous glabrous to moderately glabrous to sparsely grandiflora herbaceous holotype inner series isotypes Jansen & Harriman Kuntze lacking a cork-like lanceolate Leaf blades leaf shape lectotype Linnaeus margin and shoulders moderately ciliate moderately pilose moderately to densely morphological narrowly ovate nomen nudum oppositifolia outer series Pales papposa pappus Peduncles Petioles Phenology phyllaries ploidy Prodr R. K. Jansen radicans Ray florets Representative Specimens rooting at nodes s.n. holotype Sesse & Mocino sparsely pilose sparsely to moderately species Spilanthes Spilanthes acmella stamens Stems straight-tipped hairs stramineous strigose Stuessy subequal bristles taxa taxonomic wide yellow