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able acoustic adult alarm calls animals approach attract auditory behaviour bird song bottlenose dolphins brain breeding bullfrog bush crickets Capranica chaffinch clicks communication courtship detect dialect distance dolphins duet echolocation echolocation calls emit ewes example feeding female fight fish forest frequency modulated frequency sounds gibbons grunts hear heard high frequency human humpback whale hydrophone identify individual indri insects John Krebs Krebs laboratory lambs little brown bats locate looked loud loudspeaker low frequency male male's mammals metres mobbing moth mother neighbours nest noise Nottebohm pair pattern pitch playback experiments predators pulses recognise recorded researchers respond rival sedge warbler Seyfarth song repertoire song types sound produced sound signal sound source species spiders spinner dolphins stop singing studied swamp sparrow swim syllables syrinx target territory tree frogs ultrasonic University vervet monkeys vibration vocal vocalisations whistle white-crowned sparrow wings